Contact Info

Institution: Utah State University, Senior Partner

Phone: (435) 760-4853

Address: Department of Wildland Resources 5230 Old Main Hill, Logan, Utah 84322-5230, US


Research Disciplines

  • Biodiversity

  • Conservation Biology

  • Ecology

  • Conservation Ecology

  • Wildlife Ecology

  • Rangeland Ecology

  • Natural Resource Management

  • Grazing Management

  • Livestock and Wildlife interactions

  • Wildlife Biology

  • Zoology

  • Exotic wildlife species

  • Exotic wildlife diseases

  • Game species

  • Reintroduced wildlife species

Research Interests

bison, elk, jackrabbits, mountain goats, pikas, grazing interactions, disease ecology, exclosure experiments, rangeland ecosystem ecology


None currently


I have been a professor at USU since 2005, when I arrived as department head.  Since 2012 I have been fully engaged in research and teaching in ecology.  My research interests are diverse, but I focus mainly on the ecology of large mammals in terrestrial ecosystems.  My particular interests include interactions among species of different body size within trophic guilds, interactions between ungulates and their food plants, and differences in behavior, diet, and habitat use among sex and age classes within large mammal populations.  Together with my students and postdocs, I investigate these topics by conducting field studies on indigenous ungulates and large predators in southern African savannas and western American rangelands.  An emerging theme of my research and publications is the conservation of terrestrial ecosystem processes through the fusion of science and management.



  • 1988: PhD in animal ecology, University of the Witwatersrand, Johannesburg, South Africa
    (supervised by Norman Owen-Smith).
  • 1983: BSc Hons in zoology (1st class), University of Cape Town, South Africa.
  • 1982: BSc (distinction in zoology) University of Cape Town, South Africa.