Foundation Professor Emeritus


Contact Info

Institution: University of Nevada, Reno

Phone: (775) 230-0251

Address: NRES 1667 N. Virginia St., Reno, Nevada 89557, US


Research Disciplines

  • Avian Ecology

  • Ecology

  • Statistical Ecology

  • Wildlife Ecology

  • Population Biology

  • Livestock and Wildlife interactions

Research Interests

I have 40+ years experience working with waterfowl and 18 years experience working with sage-grouse.  This work has focued on ecological and other factors that influence demography and population dynamics.


Effects of non-native ungulates on sage-grouse


Research Scientist, U.S. Fish and Wildlife Service, 1984-85

Assistant Professor-Professor, University of Alaska Fairbanks 1985-1998

Interim Director, Institute of Arctic Biology 1998-2001

Professor, University of Nevada Reno, 2001-2019

Professor Emeritus, 2019-present


  • B.S. Electrical Engineering-University of Washington 1971
  • Ph.D. Ecology-University of California Davis 1983